Russian jets launched a second day of air strikes in Syria on Thursday, targeting areas held by an insurgent alliance that includes a group linked to al Qaeda, but not the Islamic State militants Moscow said it had hit.
Russia said it had launched eight air strikes with Sukhoi warplanes overnight, hitting four Islamic State targets. However, the areas where it said the strikes took place are not held by Islamic State.
Al-Mayadeen, a pro-Damascus television channel, said jets carried out at least 30 strikes against an insurgent alliance known as the Army of Conquest. The alliance includes the Nusra Front, al Qaeda's Syrian branch, but not Islamic State, which has declared a caliphate on swathes of Syria and Iraq.
The Army of Conquest has been advancing against government forces in northwestern Syria in recent months, and has support from regional countries that oppose both President Bashar al-Assad and Islamic State.
Russia's decision to join the war with air strikes on behalf of Assad is a major turning point in international involvement in the conflict.
The United States is leading a separate alliance waging an air war against Islamic State fighters, which means the Cold War superpower foes are now engaged in air combat over the same country for the first time since World War Two.
They say they have the same Islamic State enemies. But they also have very different friends, and opposing views of how to resolve a 4-year-old civil war that has killed more than 250,000 people and driven more than 10 million from their homes.
Washington and its allies oppose both Islamic State and Assad, believing he must leave power in any peace settlement. Moscow supports the Syrian president and believes his government should be the centerpiece of international efforts to fight extremist groups.